1.5.1 Coral reef ecosystems have the highest gross
primary productivity of all ocean areas, and the proposed area contains
several thousand square miles of coral reefs, indicating a highly
productive ecosystem. Also indicative of the area’s productivity
is the high incidence of apex predators such as sharks, jacks, and
groupers, which make up more than half of the total fish biomass in
the NWHI. A very high replacement rate of small and mid-size fish
is necessary to support an apex predator-dominated ecosystem.
1.5.2 The productivity of the proposed area can
readily be seen by comparing it to the productivity in the main Hawaiian
Islands. A comparison of both biomass and trophic structure between
reef fish communities in the NWHI and the main Hawaiian Islands showed
that across similar habitats, biomass was 260 per cent higher in the
NWHI (Friedlander and DeMartini 2002). Productivity is especially
high in the area’s inshore waters, shallow lagoons, and coral
reefs. For example, the lagoon in French Frigate Shoals produces nearly
ten times the amount of phytoplankton as produced in the same volume
of water in the open seas. The area also has extensive submerged banks,
which have high levels of primary productivity due to the existence
of expansive algal meadows. Furthermore, while apex predators represent
only three per cent of the fish biomass in the main Hawaiian Islands,
they make up 54 per cent of the biomass in the NWHI (Suthers 2004).